A Review of the Frogs of the Hyla bistincta Group   By: William E. Duellman (1930-)

Volume 15, No. 9, pp. 469 491, 4 figs. March 2, 1964

A Review of the Frogs Of the Hyla bistincta Group

BY

WILLIAM E. DUELLMAN

UNIVERSITY OF KANSAS LAWRENCE 1964

UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY

Editors: E. Raymond Hall, Chairman, Henry S. Fitch, Theodore H. Eaton, Jr.

Volume 15, No. 9, 469 491, 4 figs. Published March 2, 1964

UNIVERSITY OF KANSAS Lawrence, Kansas

PRINTED BY THE STATE PRINTER TOPEKA, KANSAS 1964

29 8590

A Review of the Frogs Of the Hyla bistincta Group

BY

WILLIAM E. DUELLMAN

CONTENTS

PAGE INTRODUCTION 471 Acknowledgments 471 THE HYLA BISTINCTA GROUP 472 Analysis of Characters 473 Key to the Species of the Hyla Bistincta Group 474 ACCOUNTS OF THE SPECIES 475 Hyla bistincta Cope 475 Hyla charadricola new species 478 Hyla robertsorum Taylor 481 Hyla pachyderma Taylor 485 Hyla crassa (Brocchi) 486 RELATIONSHIPS 489 LITERATURE CITED 491

INTRODUCTION

In the mountainous regions of Middle America there are several groups of hylid frogs that inhabit mountain streams. Some of these groups, such as Plectrohyla and Ptychohyla , have been elevated to generic rank, whereas others are retained in the large and complex genus Hyla . In the mountains of México five species of hylids that seem to compose a phyletic unit are herein referred to as the Hyla bistincta group. Since 1955 I have been accumulating specimens of, and data on, this group with the result that all specimens known to me, including the types of all named taxa, have been studied. Detailed observations have been made on the ecology and life histories of three of the species; the other two species are known to me only from preserved specimens.

Acknowledgments

For permission to examine specimens in their care I am indebted to Charles M. Bogert, American Museum of Natural History (AMNH); Doris M. Cochran, United States National Museum (USNM); Jean Guibé, Museum National d'Histoire Naturelle, Paris (MNHN); Robert F. Inger, Chicago Natural History Museum (CNHM); Hobart M. Smith, University of Illinois Museum of Natural History (UIMNH); Charles F. Walker, University of Michigan Museum of Zoology (UMMZ). (Abbreviations of institutions given above in parentheses are used throughout; the Museum of Natural History, University of Kansas is abbreviated KU.)